Phytoseiidae (Acari: Mesostigmata) in four provinces of Iran

Document Type : Original Article

Authors

1 Department of Agricultural Zoology, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organization (AREEO), Tehran, Iran

2 Department of Plant Protection, Rice Research Institute of Iran (RRII), Agricultural Research Education and Extension Organization (AREEO), Rasht, Iran

3 Plant Protection Research Department, Khorasan Razavi Agricultural & Natural Resources Research & Education Center, AREEO, Mashhad, Iran

4 University of Tehran

5 Institut Agro Montpellier (IA Montpellier), UMR CBGP INRAE / IRD / CIRAD / IA-Montpellier / Université de Montpellier, 755 Avenue du Campus Agropolis, CS 30016, 34988 Montferrier-sur-Lez cedex, France

Abstract
The present study reports the results of surveys carried out over a period of two years (2018 and 2020) in four provinces of Iran: Guilan, Razavi Khorasan, Mazandaran, and Sistan and Baluchestan. Twenty-three species from eight genera of Phytoseiidae have been collected, of which two species were newly recorded for Iran, namely: Euseius gallicus and Proprioseiopsis lineatus. Three species, namely Neoseiulus agrestis, Amblyseius obtusus, Typhlodromus (Anthoseius) haiastanius are new records for the Guilan Province. Neoseiulus californicus and Amblyseius swirskii are new records for the Mazandaran Province while Typhlodromus (Anthoseius) haiastanius is a new record for the Razavi Khorasan Province. All species collected in Sistan and Baluchestan are new records for this province. The worldwide distribution and plant hosts of all collected phytoseiids during this study are provided.

Keywords


Abbasova, E.D. (1972) Phytoseiid mites (Parasitiformes: Phytoseiidae) of Azerbaijan. Avtoreferat Dissertatsii na Soiskanie Uchenoy Stepeni Kandidata Biologicheskikh Nauk. Akadrmiya Nauk Azerbaydzhanskoy SSR, Institut Zoologii, Baku, Azerbaijan, 34 pp.
Abo-Shnaf, R.I.A. & Moraes, G.J. de. (2014) Phytoseiid mites (Acari: Phytoseiidae) from Egypt, with new records, descriptions of new species, and a key species. Zootaxa, 3865: 1–71. DOI: 10. 11646/zootaxa.3865.1.1
Amitai, S. & Grinberg, T. (1971) Description of a new phytoseiid genus and species (Acarina: Mesostigmata) from Israel. Israel Journal of Entomology, 6: 327–335.
Arutunjan, E.S. (1970) Phytoseiid mites (Phytoseiidae) on agricultural crops in the Armenian SSR. Akademii Nauk Armyanskoi SSR, Otdelenie Biologicheskikh Nauk, Dissertatsii naSoiskanie Uchenoi Stepeni Candidata Biologrcheskikh Nauk, Zooliya, 97: 31 pp.
Arutunjan, E.S. (1977) Identification manual of phytoseiid mites of agricultural crops of the Armenian SSR. Akademiya Nauk Armyanskoi SSR, Zoologicheskii Institut, Erevan, Armenia, 177 pp.
Asali Fayaz, B. & Khanjani M. (2012) Phytoseiid mites (Acari: Mesostigmata) in some regions of western and north western Iran. Journal of Crop Protection, 1(2): 161–172.
Asali Fayaz, B., Khanjani, M., Hajizadeh, J. & Ueckermann, E.A. (2012) A re-description of Typhlodromus (Anthoseius) tamaricis (Kolodochka) (Mesostigmata: Phytoseiidae), a first record for Iran. Acarologia, 52(4): 425–431.
Asali Fayaz, B., Khanjani, M., Molavi, F. & Ueckermann, E.A. (2011) Phytoseiid mites (Acari: Phytoseiidae) of apple and almond trees in regions of western and south-western Iran. Acarologia, 51(3): 371–379. DOI: 10.1051/acarologia/20112020
Asali Fayaz, B., Khanjani, M., Rahmani, H. & Ueckermann, E.A. (2017) Complementary description of Typhlodromus (Anthoseius) bagdasarjani Wainstein and Arutunjan (Acari: Mesostigmata: Phytoseiidae) based on specimens from western Iran. Acarologia, 57(2) : 255–267. DOI : 10.1051/acarologia/20164153
Athias-Henriot, C. (1959) Acariens planticoles d’Algérie. I. 5e contribution au genre Amblyseius Berlese (Phytoseiidae). II. Première liste d’Actinochitinosi (Cheyletidae, Caligonellidae, Hemi-sarcoptidae). Bulletin de l’Académie Royale de Belgique, Sciences (Ser. 5), 45: 130–-153.
Athias-Henriot, C. (1961) Mésostigmates (Uropodes exclus) édaphiques Méditerraneens (Acaromor-pha, Anactinotrichida). Acarologia, 3 : 381–509.
Athias-Henriot, C. (1962) Amblyseius swirskii, un nouveau phytoseiide voisin d'A. andersoni (Acariens anactinotriches). Annales de l'Ecole Nationale d'Agriculture d'Alger, 3 : 1–7.
Athias-Henriot, C. (1975) Nouvelles notes sur les Amblyseiini. II. Le relevé organotaxique de la face dorsale adulte (Gamasides protoadéniques, Phytoseiidae). Acarologia, 17(1) : 20–29.
Athias-Henriot, C. (1977) Nouvelles notes sur les Amblyseiini. III. Sur le genre Cydnodromus: Redé-finition, composition (Parasitiformes, Phytoseiidae). Entomophaga, 22: 61–73. DOI: 10.1007/BF 02372991
Auger, P., Tixier, M.-S., Kreiter, S. & Fauvel, G. (1999) Factors affecting ambulatory dispersal in the predaceous mite Neoseiulus californicus. Experimental and Applied Acarology, 23: 235–250. DOI: 10.1023/A:1006019014708
Basha, A.A., Yousef, A.T.A., Ibrahim, M.H. & Mostafa, E.S. (2001) Five new phytoseiids from Egypt (Acari: Gamasida: Phytoseiidae). Al-Azhar Journal of Agricultural Research, 33: 371–386.
Beaulieu, F. & Beard, J.J. (2018) Acarine biocontrol agents Neoseiulus californicus sensu Athias-Henriot (1977) and N. barkeri Hughes (Mesostigmata: Phytoseiidae) redescribed, their synonymies assessed, and the identity of N. californicus (McGregor) clarified based on examination of types. Zootaxa, 4500(4): 451–507. DOI: 10.11646/ zootaxa.4500.4.1
Beglyarov, G.A. (1958) Species of Phytoseiidae (Parasititormes: Gamasoidea) predatory upon tetranychid mites in orchards of the Krasnodar region. Trudy Vsesoiuznogo Institut Zashchity Rastenii, 10: 98–124.
Beglyarov, G.A. (1962) On the fauna of predaceous mites of the family Phytoseiidae in Krasnodar region. In: Zimina, L.S. & Shchepetilínikovoiy, V.A. (Eds.), Biologicheskiy Metod Bor'by s Vreditelyami i Boleznyami Cel'skokhozyaystvennykh kul'tur, 1: 198–203.
Beglyarov, G.A. (1981) Keys to the determination of phytoseiid mites of the USSR. Information Bulletin of the International Organization for Biological Control of Noxious Animals and Plants, East Palaearctic Section, Leningrad, Russia, 2: 97 pp.
Beglyarov, G.A. & Malov, N.A. (1977) Predaceous phytoseiid mites (Phytoseiidae: Parasitiformes) of the fauna of Moldavia and north Bukovina (Report II). Entomofagi v Zashchite Rasteniy. Vsesoyuznaya Ordena Lenina Akademiya Selískokhozyaystvennyukh Nauk Imeni V. I. Lenina. Kishinev, pp. 3–11.
Berlese, A. (1889) Acari, Myriopoda et Scorpiones hucusque in Italia reperta. Tipografia Del Seminario, 6(54): 7‌9.
Berlese, A. (1913) Systema Acarorum genera in familiis suis disposita. Acaroteca Italica, 1–2: 3–19.
Berlese, A. (1914) Acari nuovi. Manipulus IX. Redia, 10: 113–150.
Berlese, A. (1916) Centuria prima di Acari nuovi. Redia, 12: 19–‌66.
Berlese, A. (1918) Centuria quarta di Acari nuovi. Redia, 12(2): 115–192.
Blommers, L. & Chazeau J. (1974) Two new species of predator mites of the genus Amblyseius Berlese (Acarina: Phytoseiidae) from Madagascar. Zeitschrift für Angewandte Entomologie, 75: 308–315. DOI: 10.1111/j.1439-0418.1974.tb01856.x
Buitenhuis, R., Murphy, G., Shipp, L. & Scott-Dupree, C. (2015) Amblyseius swirskii in greenhouse production systems: a floriculture perspective. Experimental and Applied Acarology, 65: 451–464. DOI: 10.1007/s10493-014-9869-9
Byng, J.W., Smets, E.F., van Vugt, R., Bidault, E., Davidson, C., Kenicer, G., Chase, M.W. & Christenhusz, M.J.M. (2018) The phylogeny of angiosperms poster: a visual summary of APG IV family relationships and floral diversity. The Global Flora, p. 4‌7.
Calvo, F.J., Knapp, M., van Houten, Y.M., Hoogerbrugge, H. & Belda, J. E. (2015) Amblyseius swirskii: What made this predatory mite such a successful biocontrol agent? Experimental and Applied Acarology, 65: 419–433. DOI: 10.1007/s10493-014-9873-0
Castagnoli, M. & Simoni, S. (2003) Neoseiulus californicus (McGregor) (Acari Phytoseiidae): Survey of biological and behavioural traits of a versatile predator. Redia, 86: 153–164.
Chant, D.A. (1957) Descriptions of some phytoseiid mites (Acarina, Phytoseiidae). Part I. Nine new species from British Columbia with keys to the species of British Columbia. Part II. Redescriptions of eight species described by Berlese. The Canadian Entomologist, 89(7): 289–308. DOI: 10.4039/Ent89289-7
Chant, D.A. (1959) Phytoseiid mites (Acarina: Phytoseiidae). Part I. Bionomics of seven species in southeastern England. Part II. A taxonomic review of the family Phytoseiidae, with descriptions of thirty-eight new species. The Canadian Entomologist, 61(12): 1–166. DOI: 10.4039/entm9112 fv
Chant, D.A. & Athias-Henriot, C. (1960) The genus Phytoseius Ribaga, 1902 (Acarina: Phytoseiidae). Entomophaga, 5: 213–228. DOI: 10.1007/BF02383272
Chant, D.A. & Hansell, R.I.C. (1971) The genus Amblyseius (Acarina: Phytoseiidae) in Canada and Alaska. Canadian Journal of Zoology, 49(5): 703–758. DOI: 10.1139/z71-110
Chant, D.A., Hansell, R.I.C. & Rowell, H. (1978) A numerical taxonomic study of variation in populations of Typhlodromus cuudiglans Schuster (Acarina: Phytoseiidae). Canadian Journal of Zoology, 56: 55–65. DOI: 10.1139/z78-007
Chant, D.A., Hansell, R.I.C. & Yoshida-Shaul, E. (1974) The genus Typhlodromus Scheuten (Acarina: Phytoseiidae) in Canada and Alaska. Canadian Journal of Zoology, 52: 1265–1291. DOI: 10.1139/z74-168
Chant, D.A. & McMurtry, J.A. (1994) A review of the subfamilies Phytoseiinae and Typhlodrominae (Acari: Phytoseiidae). International Journal of Acarology, 20(4): 223‌–310. DOI: 10.1080/01647 959408684022
Chant, D.A. & McMurtry, J.A. (2003) A review of the subfamily Amblyseiinae Muma (Acari: Phytoseiidae): Part I. Neoseiulini new tribe. International Journal of Acarology, 29(1): 3‌–46. DOI: 10.1080/01647950308684319
Chant, D.A. & McMurtry, J.A. (2004) A review of the subfamily Amblyseiinae Muma (Acari: Phytoseiidae): Part III. The tribe Amblyseiini Wainstein, subtribe Amblyseiina n. subtribe. International Journal of Acarology, 30(3): 171–228. DOI: 10.1080/01647950408684388
Chant, D.A. & McMurtry, J.A. (2005a) A review of the subfamily Amblyseiina Muma (Acari: Phytoseiidae): Part V. Tribe Amblyseiini, subtribe Proprioseiopsina Chant & McMurtry. Interna-tional Journal of Acarology, 31(1): 3–22. DOI: 10.1080/01647950508684412
Chant, D.A. & McMurtry, J.A. (2005b) A review of the subfamily Amblyseiinae Muma (Acari: Phytoseiidae) Part VI. The tribe Euseiini n. tribe, subtribes Typhlodromalina n. subtribe, Euseiina n. subtribe, and Ricoseiina n. subtribe. International Journal of Acarology, 31(3): 187‌–224. DOI: 10.1080/01647950508684424
Chant, D.A. & McMurtry, J.A. (2007) Illustrated keys and diognoses for the genera and subgenera of the Phytoseiidae of the world (Acari: Mesostigmata). Indira Publishing House, West Bloomfield, 219 pp.
Chant, D.A. & Yoshida-Shaul, E. (1989) A world review of the tiliarum species group in the genus Typhlodromus Scheuten (Acari: Phytoseiidae). Canadian Journal of Zoology, 67(4): 1006–1046.
Chant, D.A. & Yoshida-Shaul, E. (1991) Adult ventral setal patterns in the family Phytoseiidae (Acari: Gamasina). International Journal of Acarology, 17(3): 187–199. DOI: 10.1080/0164795 9108683906
Chaudhri, W.M., Akbar, S. & Rasool, A. (1979) Studies on the predatory leaf inhabiting mites of Pakistan. University of Agriculture, Faisalabad, Pakistan, 243 pp.
Çobanoğlu, S., Akçakoyunlóuoğlu, K. & Çalmaşur, Ö. (2020) Typhlodromus (Anthoseius) caudiglans (Schuster, 1959) (Acari: Phytoseiidae) as predatory mite in Erzurum-Turkey. Tarim Bilimleri Dergisi, 26: 130–137. DOI: 10.15832/ankutbd.498043
Congdon, B.D. (2002) The family Phytoseiidae (Acari) in western Washington State with descriptions of three new species. International Journal of Acarology, 28(1): 3–‌27. DOI: 10.1080 /01647950208684275
Corpuz, L.A. & Rimando, L. (1966) Some Philippine Amblyseiinae (Phytoseiidae: Acarina). Philippines Agricullture, 50: 114–136.
Cunliffe, F. & Baker, E.W. (1953) A guide to the predatory phytoseiid mites of the United States. Pinellas Biology Laboratory, Inc., USA, 1, 28 pp.
Daneshvar, H. (1987) Some predacious mites from Iran, with descriptions of one new genus and six new species (Acari: Phytoseiidae, Ascidae). Entomologie et Phytopathologie Appliquées, 54 (1–2): 13–37 (In English), 55–73 (In Persian).
Daneshvar, H. & Denmark, H.A. (1982) Phytoseiids of Iran (Acarina: Phytoseiidae). International Journal of Acarology, 8: 3–14. DOI: 10.1080/01647958208683272
De Leon, D. (1957) Three new Typhlodromus from southern Florida (Acarina: Phtyoseiidae). The Florida Entomologist, 40: 141–144. DOI: 10.2307/3492041
De Leon, D. (1959) Two new genera of phytoseiid mites with a note on Proprioseius meridionalis Chant (Acarina: Phytoseiidae). Entomological News, 70(10): 257–262.
De Leon, D. (1962) Twenty-three new phytoseiids, mostly from southeastern United States (Acarina: Phytoseiidae). The Florida Entomologist, 45(1): 11–27. DOI: 10.2307/3492899
De Leon, D. (1966) Phytoseiidae of British Guyana with keys to species (Acarina: Mesostigmata). Studies on the Fauna of Suriname and Other Guyanas, 8: 81–102.
Demite, P.R., McMurtry, J.A. & Moraes, G.J. de (2014) Phytoseiidae Database: a website for taxonomic and distributional information on phytoseiid mites (Acari). Zootaxa, 3795(5): 571–577. DOI: 10.11646/zootaxa.3795.5.6
Demite, P.R., Moraes, G.J. de, McMurtry, J.A., Denmark, H.A. & Castilho, R.C. (2022) Phytoseiidae database. University of São Paulo, Brazil. Available from: www.lea.esalq.usp.br/phytoseiidae (Accessed on September 2022).
Denmark, H.A. (1965) Four new Phytoseiidae (Acari: Mesostigmata) from Florida. The Florida Entomologist, 48: 89–95. DOI: 10.2307/3493097
Denmark, H.A. (1966) Revision of the genus Phytoseius Ribaga, 1904 (Acarina: Phytoseiidae). Florida Department of Agriculture Bulletin, 6: 1–105.
Denmark, H.A. & Evans, G.A. (2011) Phytoseiidae of North America and Hawaii (Acari: Mesostigmata). Indira Publishing House, West Bloomfield, USA, 451 pp.
Denmark, H.A. & Muma, M.H. (1989) A revision of the genus Amblyseius Berlese (1914) (Acari: Phytoseiidae). Occasional papers of the Florida State Collection of Arthropods (USA), 4: 1–149.
Denmark, H.A. & Welbourn, W.C. (2002) Revision of the genera Amblydromella, Muma and Anthoseius, De Leon (Acari: Phytoseiidae). International Journal of Acarology, 28(4): 291–316. DOI: 10.1080/01647950208684308
Döker, I., Kazak, C. & Karut, K. (2020) The genus Amblyseius Berlese (Acari: Phytoseiidae) in Turkey with discussion on the identity of Amblyseius meridionalis. Systmatic and Applied Acarology, 25(8): 1395‌–1420. DOI: 10.11158/saa.25.8.4
Döker, I., Witters, J., Pijnakker, J., Kazak, C., Tixier, M.-S. & Kreiter, S. (2014) Euseius gallicus Kreiter and Tixier (Acari: Phytoseiidae) is present in four more countries in Europe: Belgium, Germany, The Netherlands and Turkey. Acarologia, 54(3): 245–248. DOI: 10.1051/acarologia/2 0142132
Dosse, G. (1957) Morphologie und biologie von Typhlodromus zwoelferi n. sp. (Acar., Phytoseiidae). Zeitschrift fur Angewandte Entomologie, 41(2–3): 301–311. DOI: 10.1111/j.1439-0418.1957.tb0 1295.x
Dosse, G. (1958) Uber einige neue Raubmilbenarten (Acari: Phytoseiidae). Pflanzenschutz Berichte, 21 : 44–61
Dosse, G. (1967) Schadmilben des Libanons und ihre Pradatoren. Zeitschrift fur Angewandte Entomologie, 59: 16–48. DOI: 10.1111/j.1439-0418.1967.tb03837.x
Duso, C. & Fontana, P. (2002) On the identity of Phytoseius plumifer (Canestrini & Fanzago, 1876) (Acari: Phytoseiidae). Acarologia, 42(2): 127–136.
Duso, C. & Moretto, S. (1994) Osservazioni preliminari sui comportamento dell’acaro predatore Phytoseius plumifer (Can. et Fanz.). Memorie della Società Entomologica Italiana, 72: 533–540.
Easterbrook, M.A., Fitzgerald, J.D. &, Solomon, M.G. (2001) Biological control of strawberry tarsonemid mite Phytonemus pallidus and two-spotted spider mite Tetranychus urticae on strawberry in the UK using species of Neoseiulus (Amblyseius) (Acari: Phytoseiidae). Experimental Applied Acarology, 25: 25–36. DOI: 10.1023/A:1010685903130
Ehara, S. (1966) A tentative catalogue of predatory mites of Phytoseiidae known from Asia, with descriptions of five new species from Japan. Mushi, 39: 9‌–30.
Ehara, S. (1985) Five species of phytoseiid mites from Japan with descriptions of two new species (Acarina, Phytoseiidae). Zoological Science, 2(1): 115–121.
Ehara, S. & Amano, H. (1998) A revision of the mite family Phytoseiidae in Japan (Acari: Gamasina), with remarks on its biology. Species Diversity, 3(1): 25–73. DOI: 10.12782/specdiv.3.25
Ehara, S. & Amano, H. (2004) Checklist and keys to Japanese, Amblyseiinae (Acari: Gamasina: Phytoseiidae). Journal Acarology Society Japan, 13(1): 1–30. DOI: 10.2300/acari.13.1
Ehara, S. & Bhandhufalck, A. (1977) Phytoseiid mites of Thailand (Acarina: Mesotigmata). Journal of the Faculty of Education of Tottori University, Natural Sciences, 27(2): 43–82.
Ehara, S., Okada, Y. & Kato, H. (1994) Contribution to the knowledge of the mite family Phytoseiidae in Japan (Acari: Gamasina). Journal of the Faculty of Education of Tottori University, Natural Sciences, 42(2): 119–160.
El-Badry, E.A. (1967) Five new phytoseiid mites from U. A. R., with collection notes on three other species (Acarina: Phytoseiidae). Indian Journal of Entomology, 29: 177–184.
Evans, G.O. (1965) The ontogenetic development of the chaetotaxy of the tarsi of legs II-IV in the Antennophorina (Acari: Mesostigmata). Annual Magasin of Natural History, Series 13, 8: 81–83. DOI: 10.1080/00222936508651543
Evans, G.O. & Edland, T. (1998) The genus Anthoseius De Leon (Acari: Mesostigmata) in Norway. Fauna Norvegica, Ser. B, 45: 41–62.
Famah Sourassou, N., Hanna, R., Zannou, I., Moraes, G.J. de, Negloh, K. & Sabelis, M.W. (2011) Morphological variation and reproductive incompatibility of three coconut-mite-associated populations of predatory mites identified as Neoseiulus paspalivorus (Acari: Phytoseiidae). Experimental and Applied Acarology, 53: 323–338. DOI: 10.1007/s10493-010-9413-5
Faraji, F., Çobanoğlu, S. & Çakmak, I. (2011) A checklist and a key for the Phytoseiidae species of Turkey with two new species records (Acari: Mesostigmata). International Journal of Acarology, 37(Supl. 1): 221–243. DOI: 10.1080/01647954.2011.558851
Faraji, F., Hajizadeh, J., Ueckermann, E.A., Kamali, K. & McMurtry, J.A. (2007) Two new records for Iranian phytoseiid mites with synonymy and keys to the species of Typhloseiulus Chant and McMurtry and Phytoseiidae in Iran (Acari: Mesostigmata). International Journal of Acarology, 33(3): 231–239. DOI: 10.1080/01647950708684527
Faraji, F., Sakenin-Chelav, H., Kamali, K. & McMurtry, J.A. (2008) Four new species records of Phytoseiidae (Acari: Mesostigmata) for Iran, and description of variability in the spermatheca of Typhlodromus bakeri. Systematic and Applied Acarology, 13: 123–132.
Ferragut, F. (2018) New records of phytoseiid mites of the subfamilies Typhlodrominae and Phytoseiinae (Acari: Phytoseiidae) from Spain, with description of a new species and re-description of four species of Typhlodromus Scheuten. Systematic and Applied Acarology, 23(5): 883–910. DOI: 10.11158/saa.23.5.8
Ferragut, F., Pérez Moreno, I., Iraola, V. & Escudero, A. (2010) Ácaros depredadores em las plantas cultivadas. Família Phytoseiidae. Ediciones Agrotécnicas, Madrid, 202 pp.
Garman, P. (1948) Mite species from apple trees in Connecticut. Connecticut Agricultural Experi-ment Station, Bulletin, New Haven, USA, 520: 1–27.
Garman, P. (1958) New species belonging to the genera Amblyseius and Amblyseiopsis, with keys to Amblyseius, Amblyseiopsis, and Phytoseiulus. Annual Entomological Society of America, 51: 69‌–79. DOI: 10.1093/aesa/51.1.69
Gerson, U., Smiley, R.L. & Ochoa, R. (2003) Mites (Acari) for Pest Control. Blackwell Science Ltd., UK, 539 pp. DOI: 10.1002/9780470750995
Ghai, S. & Gupta, S.K. (1984) A new species of Treatia Krantz and Khot (Acari: Otophe Idomenidae [Otopheidomenidae]) with a new record of Amblyseius Berlese (Acari: Phytoseiidae) from India. Bulletin of the Zoological Survey of India, 6(1–3): 171–175.
Gomelauri, L.A. (1968) Three new species of mites of the family Phytoseiidae, in southern Georgia. Bulletin of the Academy of Sciences of the Georgian SSR, Zoology and Parasitology, 52(2): 515–520.
Gomez-Moya, C.A. & Ferragut, F. (2009) Spatial distribution pattern and efficacy of Neoseiulus californicus and Phytoseiulus persimilis (Acari: Phytoseiidae) in the control of red spider mites on vegetables under semi-field conditions. Bolletin Sanidad Vegetal Plagas, 35: 377–390.
Guanilo, A.D., Moraes, G.J. de & Knapp, M. (2008a) Phytoseiid mites (Acari: Phytoseiidae) of the subfamily Amblyseiinae Muma from Peru, with description of four new species. Zootaxa, 1880: 1–47. DOI: 10.11646/zootaxa.1880.1.1
Guanilo, A.D., Moraes, G.J. de, Toledo, S. & Knapp, M. (2008b) Phytoseiid mites (Acari: Phytosei-idae) from Argentina, with description of a new species. Zootaxa, 1884: 1–35. DOI: 10.11646/ zootaxa.1884.1.1
Gupta, S.K. (1986) Fauna of India. Family Phytoseiidae. Zoological Survey of India, Calcutta, India, 350 pp.
Hajizadeh, J. & Faraji, F. (2016) Identification guide and diagnosis, key for predatory mites, of the family Phytoseiidae of Iran. University of Guilan Press, Rasht, Iran, 164 pp. (In Persian).
Hajizadeh, J., Hosseini, R. & McMurtry, J.A. (2002) Phytoseiid mites (Acari: Phytoseiidae) associated, with eriophyid mites (Acari: Eriophyidae) in Guilan province of Iran. International Journal of Acarology, 28(4): 373–378. DOI: 10.1080/01647950208684313
Hajizadeh, J. & Mortazavi, S. (2015) The genus Euseius Wainstein (Acari: Phytoseiidae) in Iran, with a revised key to Iranian phytoseiid mites. International Journal of Acarology, 41(1): 53–66.
Hajizadeh, J. & Nazari, M. (2012) A checklist and key for the phytoseiid mites (Acari: Phytoseiidae) of citrus orchards in Iran, with a new record for Iranian phytoseiid mites. Systematic and Applied Acarology, 17: 388–396. DOI: 10.11158/saa.17.4.7
Hirschmann, W. (1962) Gangystematik der Parasitiformes. Acarologie, Schriftenreihe fur Verglei-chende, Milbenkunde. Hirschmann-Verlag, Furth/Bay, 5(5–6): 80 pp. + 32 plates.
Hughes, A.M. (1948) The mites associated with stored food products. Ministry of Agriculture and Fisheries, H. M. Stationary Office, London, 168 pp.
Hughes, A.M. (1961) The mites of stored food. Ministry of Agriculture, Fishery, and Food. Technical Bulletin, First Edition, London, 9: 287 pp.
Jafari, S., Fathipour, Y. & Faraji, F. (2011) Redescriptions of Amblyseius meghriensis Arutunjan and Typhlodromus haiastanius (Arutunjan) with discussion on using preanal pores as a character in the subgenus, Anthoseius (Mesostigmata: Phytoseiidae). International Journal of Acarology, 37 (3): 244–254. DOI: 10.1080/01647954.2010.514288
Karg, W. (1960) Zur Kenntnis der Typhlodromiden (Acarina, Parasitiformes) aus Acker- und Grün-landböden. Zeitschrift fur Angewandte Entomologie, 47: 440–452. DOI: 10.1111/j.1439-0418. 1960.tb02848.x
Karg, W. (1970) Neue Arten der Raubmilbenfamilie Phytoseiidae Berlese, 1916 (Acarina: Parasiti-formes). Deutsche Entomologische Zeitschrift N. F., 17: 289–301. DOI: 10.1002/mmnd.4810170 402
Karg, W. (1971) Acari (Acarina), Milben, Unterordnung Anactinochaeta (Parasitiformes): Die freilebenden Gamasina (Gamasides), Raubmilben. Die Tierwelt Deutschland und der angrenzenden Meeresteile, 59. Teil, VEB Gustav Fischer Verlag, Jena, Germany, 475 pp.
Karg, W. (1975) Nutzlingsschonende Spinnmilbenbekampfung im Obstbau. Gartenbau, 22: 51–54.
Karg, W. (1977) Zur Kenntnis der Gattung Amblyseius Berlese, 1904. Abhandlungen und Berichte des Naturkundemuseums Görlitz, 51(6): 1–9.
Karg, W. (1982) Diagnostic and systematics of predatory mites of the family Phytoseiidae Berlese in orchards. Zoologische Jahrbücher Abteilung für Systematik, Geographie und Biologie der Tiere, 109: 188–210.
Karg, W. (1983) Systematische untersuchung der Gattungen und Untergattungen der Raubmilben familie Phytoseiidae Berlese, 1916, mit der beschreibung von 8 neuen Arten. Mitteilungen aus dem Zoologischen Museum in Berlin, 59(2): 293–328. DOI: 10.1002/mmnz.4830590203
Karg, W. (1989) Neue Raubmilbenarten der Gattuig Proprioseiopsis Muma, 1961 (Acarina, Parasiti-formes) mit Bestimmungsschlusseln. Zoologische Jahrbücher Abteilung für Systematik, Geogra-phie und Biologie der Tiere, 116(2): 199–216.
Karg, W. (1991) Die Raubmilbenarten der Phytoseiidae Berlese (Acarina) Mitteleuropas sowie angrenzender Gebiete. Zoologische Jahrbücher Abteilung für Systematik, Geographie und Biologie der Tiere, 118(1): 1–64.
Karg, W. (1993) Acari (Acarina), Milben Parasitiformes (Anactinochaeta) Cohors Gamasina Leach. Raubmilbe. Die Tierwelt Deutschlands: 59, Second Edition, Gustav Fischer Verlag, Jena, Germany, 523 pp.
Kasap, I. & Çobanoğlu, S. (2009) Phytoseiid mites of Hakkâri province, with Typhlodromus (Anthoseius) tamaricis Kolodochka, 1982 (Acari: Phytoseiidae), a new record for the predatory mite fauna of Turkey. Türk Zooloji Dergisi, 33: 301–308. DOI: 10.3906/zoo-0805-9
Kazemi, S., Mohammad-Doustaresharaf, M. & Döker, I. (2022) An annotated checklist of the Iranian Phytoseiidae (Acari: Mesostigmata), with an updated key to the species. Systematic and Applied Acarology, 27(4): 697–748. DOI: 10.11158/saa.27.4.6
Koch, C.L. (1839) Deutschlands Crustaceen, Myriapoden und Arachniden. Regensburg, Germany, 5–6(25): 22; 5–6 (27): 6, 13.
Kolodochka, L.A. (1973) Predaceous phytoseiid mites (Parasitiformes: Phytoseiidae) from the forest steppe of the Ukrainian SSR. Part I. Species of the genus Amblyseius. Vestnik Zoologii, 5: 78–81.
Kolodochka, L.A. (1982) New phytoseiid mites (Parasitiformes: Phytoseiidae) from Turkmen. Vestnik Zoologii, 6: 7–13.
Kolodochka, L.A. (1998) Two new tribes and the main results of a revision of Paleartic phytoseiid mites (Parasitiformes, Phytoseiidae) with the family system concept. Vestnik Zoologii, 32(1–2): 51–63.
Kolodochka, L.A. (2006) Phytoseiid mites of the Palaearctic Region (Parasitiformes, Phytoseiidae): faunistic, taxonomy, ecomorphology, evolution. Vestnik Zoologii, 21: 1–250.
Kolodochka, L.A. (2018) Two new species of the genus Neoseiulus (Parasitiformes, Phytoseiidae) with redescriptions of N. bicaudus and N. micmac based on holotypes. Vestnik Zoologii, 52(4): 295–306. DOI: 10.2478/vzoo-2018-0031
Kreiter, S., Amiri, K., Douin, M., Bohinc, T., Trdan, S. & Tixier, M.-S. (2020a) Phytoseiid mites of Slovenia (Acari: Mesostigmata): new records and first description of the male of Amblyseius microorientalis. Acarologia, 60(2): 203–242. DOI: 10.24349/acarologia/20204364
Kreiter, S., Payet, R.-M., Douin, M., Fontaine, O., Fillâtre, J. & Le Bellec, F. (2020b) Phytoseiidae of La Réunion Island (Acari: Mesostigmata): three new species and two males described, new synonymies, and new records. Acarologia, 60(1): 111–195. DOI: 10.24349/acarologia/20184273
Kreiter, S., Vicente, V., Tixier, M.-S. & Fontaine, O. (2016a) An unexpected occurrence of Amblyseius swirskii Athias-Henriot in La Reunion Island (Acari: Phytoseiidae). Acarologia, 56(2): 175–181. DOI: 10.1051/acarologia/20162254
Kreiter, S., Vicente, V., Tixier, M.-S., Fontaine, O., Avril, I., Cottineau, J.-S. & Fillâtre, J. (2016b) Présence inattendue d’Amblyseius swirskii Athias-Henriot à l’Île de La Réunion. Phytoma-La santé des végétaux, 695: 38–42.
Liao, J.-R., Ho, C.-C. & Ko, C.-C. (2021b) Survey of phytoseiid mites (Acari: Mesostigmata) in the Penghu Islands with two new records and descriptions of two new species. Systematic and Applied Acarology, 26(4): 641–671. DOI: 10.11158/saa.26.4.1
Lindquist, E.E. (1994) Some observations on the chaetotaxy of the caudal body region of gamasine mites (Acari: Mesostigmata), with a modified notation for some ventrolateral body setae. Acarologia, 35: 323–‌326.
Lindquist, E.E. & Evans, G.W. (1965) Taxonomic concepts in the Ascidae, with a modified setal nomenclature for the idiosoma of the Gamasina. Memoirs of the Entomological Society of Canada, 47: 55–66. DOI: 10.4039/entm9747fv
Louni, M., Jafari, S. & Shakarami, J. (2016) The effect of temperature on the functional response and prey consumption of Phytoseius plumifer (Phytoseiidae) fed on Rhyncaphytoptus ficifoliae (Diptilomiopidae). Plant Pest Research, 6(1): 71–83.
McGregor, E.A. (1954) Two new mites in the genus Typhlodromus (Acarina: Phytoseiidae). Southern California Academy of Science Bulletin, 53: 89–92.
Meshkov, Yu.I. (1991) Euseius amissibilis sp. n. (Parasitiformes, Phytoseiidae), a new species of mite from Tadjikistan. Zoologicheskii Zhurnal, 70: 138–140.
Moraes, G.J. de, McMurtry, J.A. & Yaninek, J.S. (1989) Some phytoseiid mites (Acari, Phytoseiidae) from tropical Africa with description of a new species. International Journal of Acarology, 15(2): 95‌–102. DOI: 10.1080/01647958908683830
Moraes, G.J. de, McMurtry, J.A. & Denmark, H.A. (1986) A catalog of the mite family Phytoseiidae. References to taxonomy, synonymy, distribution and habitat. EMBRAPA-DDT, Brasilia, Brazil, 353 pp.
Moraes, G.J. de, McMurtry, J.A., Denmark, H.A. & Campos, C.B. (2004) A revised catalog of the mite family Phytoseiidae. Zootaxa, 434: 1–494. DOI: 10.11646/zootaxa.434.1.1
Moraes, G.J. de, Kreiter, S. & Lofego, A.C. (2000) Plant mites (Acari) of the French Antilles. 3. Phytoseiidae (Gamasida). Acarologia, 40(3): 237–‌264.
Muma, M.H. (1961) Subfamiles, genera, and species of Phytoseiidae (Acarina: Mesostigmata). Florida State Museum Bulletin, USA, 5(7): 267–302.
Muma, M.H. (1967) New Phytoseiidae (Acarina: Mesostigmata) from southern Asia. The Florida Entomologist, 50: 267–280. DOI: 10.2307/3493156
Muma, M.H. & Denmark, H.A. (1970) Phytoseiidae of Florida. Arthropods of Florida and neighboring land areas, 6. Florida Department of Agriculture and Consumer Services, Division of Plant Industry, Gainesville, USA, 150 pp.
Nazari, M., Hajizadeh J. & Noei, J. (2011) Twelve species of Phytoseiidae (Acari: Mesostigmata) from citrus orchards of Guilan Province, Iran. In: Kazemi, S. & Saboori, A. (Eds.), Proceedings of the First Persian Congress of Acarology. 22–23 December 2011, International Center for Science, High Technology & Environmental Sciences, Kerman, Iran, 34 pp.
Nesbitt, H.H.J. (1951) A taxonomic study of the Phytoseiinae (Family Laelaptidae) predaceous upon Tetranychidae of economic importance. Zoologische Verhandelingen, 12: 1–96.
Okassa, M., Tixier, M.-S., Cheval, B. & Kreiter, S. (2009) Molecular and morphological evidence for a new species status within the genus Euseius (Acari: Phytoseiidae). Canadian Journal of Zoology, 87: 689–698. DOI: 10.1139/Z09-057
Oliveira, D.C., Charanasri, V., Kongchuensin, M., Konvipasruang, P., Chandrapatya, A. & Moraes G.J. de (2012) Phytoseiidae of Thailand (Acari: Mesostigmata), with a key for their identify-cation. Zootaxa, 3453: 1–24. DOI: 10.11646/zootaxa.3453.1.1
Ostovan, H., Faraji, F., Kamyab, F. & Khadempour F. (2012) Notes on Neoseiulus paspalivorus (De Leon) and Proprioseiopsis messor (Wainstein) (Acari: Phytoseiidae) collected in Iran. Acarologia, 52(1): 51–58. DOI: 10.1051/acarologia/20122032
Oudemans, A.C. (1905) Acarologische Aanteekeningen, XIX. Entomologische Berichten, 2(25): 4–12.
Panahi, L., H., Askarianzadeh, A. & Jalaeian, M. (2014) Phytoseiid mites (Acari: Phytoseiidae) of fruit orchards in cold regions of Razavi Khorasan province (northeast Iran), with redescription of two species. Persian Journal of Acarology, 3(1): 27–40.
Prasad, V. (1968) Amblyseius mites from Hawaii. Annals of the Entomological Society of America, 61(6): 1514–1521. DOI: 10.1093/aesa/61.6.1514
Pritchard, A.E. & Baker, E.W. (1962) Mites of the family Phytoseiidae from Central Africa, with remarks on the genera of the world. Hilgardia, 33(7): 205–309. DOI: 10.3733/hilg.v33n07p205
Ragusa, S. & Athias-Henriot, C. (1983) Observations on the genus Neoseiulus Hughes (Parasiti-formes, Phytoseiidae). Redefinition. Composition. Geography. Description of two new species. Revue Suisse de Zoologie, 90(3): 657–678. DOI: 10.5962/bhl.part.82005
Rahmani, H., Kamali, K. & Faraji, F. (2010) Predatory mite fauna of Phytoseiidae of northwest Iran (Acari: Mesostigmata). Turkish Journal of Zoology, 34(4): 497–508. DOI: 10.3906/zoo-0902-23
Rasmy, A. & El-Banhawy, E.M. (1974a) Behaviour and bionomics of the predatory mite, Phytoseius plumifer (Acarina: Phytoseiidae) as affected by physical surface features of host plants. Entomophaga, 19: 255–257. DOI: 10.1007/BF02371050
Rasmy, A. & El-Banhawy, E.M. (1974b) The phytoseiid mite Phytoseius plumifer as a predator of the eriophyid mite Aceria fiats (Acarina). Entomophaga, 19(3): 255–257. DOI: 10.1007/BF0237 1050
Rasmy, A. & El-Banhawy, E.M. (1975) Biology and predatory efficiency of two phytoseiid mites as affected by long term pollen feeding. Entomophaga, 20(1): 93–95. DOI: 10.1007/BF02373454
Rhodes, L.M. & Liburd, O.E. (2006) Evaluation of predatory mites and Acramite for control of twospotted spider mites in strawberries in north central Florida. Journal of Economic Entomology, 99(4): 1291–1298. DOI: 10.1093/jee/99.4.1291
Ribaga, C. (1904) Gamasidi planticoli. Rivista Patologia Vegetale, 10: 175–178.
Rivnay, T. & Swirski, E. (1980) Four new species of phytoseiid mites (Acarina: Mesostigmata) from Israel. Phytoparasitica, 8: 173–187. DOI: 10.1007/BF03158314
Rodriguez-Reina, J.M., Garcia-Mari, F. & Ferragut, F. (1992) Predatory activity of phytoseiid mites on different developmental stages of the Western flower thrips Frankliniella occidentalis. Bolletin Sanidad Vegetal Plagas, 18: 253–263.
Rowell, H.J., Chant, D.A. & Hansell, R.I.C. (1978) The determination of setal homologies and setal patterns on the dorsal shield in the family Phytoseiidae. Canadian Entomologist, 110: 859–‌876. DOI: 10.4039/Ent110859-8
Ryu, M.O., Lee, W.K. & Cho, S.R. (2001) Phytoseiid mites (Acari: Phytoseiidae) from the pear field of Naju District in Korea. Korean Journal of Soil Zoology, 6(1–2): 7–9.
Salmane, I. (1999) Soil free-living predatory Gamasina mites (Acari, Mesostigmata) from the coastal meadows of Riga Gulf, Latvia. Latvijas Entomologs, 37: 104–114.
Scheuten, A. (1857) Einiges uber Milben. Archiv für Naturgeschichte, 23: 104–112.
Schicha, E. (1978) Two new Typhlodromus from Australia and T. caudiglans Schuster redescribed (Acari: Phytoseiidae). Acarologia, 10(3): 317–326.
Schicha, E. (1979) Three new species of Amblyseius Berlese (Acarina: Phytoseiidae) from Australia. Proceedings of the Linnean Society of New South Wales, 103: 217–226.
Schicha, E. (1981) A new species of Amblyseius (Acari: Phytoseiidae) from Australia compared with ten closely related species from Asia, America and Africa. International Journal of Acarology, 7: 203–216. DOI: 10.1080/01647958108683262
Schicha, E. (1987) Phytoseiidae of Australia and neighboring areas. Indira Publishing House, West Bloomfield, Michigan, USA, 187 pp.
Schicha, E. & Corpuz-Raros, L.A. (1992) Phytoseiidae of the Philippines. Indira Publishing House, West Bloomfield, Michigan, USA, 190 pp.
Schuster, R.O. (1959) A new species of Typhlodromus near T. bakeri (Garman) and a consideration of the occurrence of T. rhenanus in California. Proceedings of the Entomological Society of Washington, 61(2): 88–90.
Schuster, R.O. & Pritchard, A.E. (1963) Phytoseiid mites of California. Hilgardia, 34: 191–285. DOI: 10.3733/hilg.v34n07p191
Specht, H.B. (1968) Phytoseiidae (Acarina: Mesostigmata) in the New Jersey apple orchard environ-ment with descriptions of spermathecae and three new species. Canadian Entomologist, 100: 673–692. DOI: 10.4039/Ent100673-7
Stathakis, T.I., Kapaxidi, E.V. & Papadoulis, G.T. (2016) A new species and three new records of Phytoseiidae (Acari: Mesostigmata) found on coastal and wetland vegetation in Greece. Systematic and Applied Acarology, 21: 567–582. DOI: 10.11158/saa.21.5.2
Szabó, A., Gál, C. & Pénzes, B. (2013) The occurrence of predatory mites in the Kunsa′g wine region in Hungary. Növvéd, 49: 193–197.
Tixier, M.-S., Auger, P., Migeon, A., Douin, M., Fossoud, A., Navajas, M. & Arabuli, T. (2021) Integrated taxonomy supports the identification of some species of Phytoseiidae (Acari: Mesostigmata) from Georgia. Acarologia, 61(4): 824–844. DOI: 10.24349/m2Rp-WodG
Tixier, M.-S., Guichou, S. & Kreiter, S. (2008) Morphological variation in the biological control agent Neoseiulus californicus (McGregor) (Acari: Phytoseiidae): consequences for diagnostic reliability and synonymies. Invertebrate Systematics, 22: 453–469. DOI: 10.1071/IS07052
Tixier, M.-S., Kreiter, S., Okassa, M. & Cheval, B. (2010) A new species of the genus Euseius Wainstein (Acari: Phytoseiidae) from France. Journal Natural History, 44(3–4): 241–254. DOI: 10.1080/00222930903383529
Tixier, M.-S., Lopes, I., Blanc, G., Dedieu, J.L. & Kreiter S. (2014) Phytoseiid mite diversity (Acari: Mesostigmata) and assessment of their spatial distribution in French apple orchards. Acarologia, 54(1): 97–111. DOI: 10.1051/acarologia/20142114
Tixier, M.-S., Tsolakis, H., Ragusa, S., Poiso, A., Ferrero, M., Okassa, M. & Kreiter, S. (2011) Integrative taxonomy demonstrates the unexpected synonymy between two predatory mite species: Cydnodromus idaeus and C. picanus (Acari: Phytoseiidae). Invertebrate Systematics, 25: 273–281.
Tixier, M.-S., dos Santos Vicente, V., Douin, M., Duso, C. & Kreiter, S. (2017) Great molecular variation within the species Phytoseius finitimus (Acari: Phytoseiidae): implications for diagnosis decision within the mite family Phytoseiidae. Acarologia, 57(3): 493–515. DOI: 10.24349/ acarologia/20174168
Ueckermann, E.A. & Loots, G.C. (1988) The African species of the subgenera Anthoseius De Leon and Amblyseius Berlese (Acari: Phytoseiidae). Entomological Memoirs, Department of Agricul-ture and Water Supply, Republic of South Africa, 73: 168 pp.
van der Merwe, G.G. (1965) South African Phytoseiidae (Acarina). I. Nine new species of the genus Amblyseius Berlese. Journal of the Entomological Society of South Africa, 28: 57‌–76.
van der Merwe, G.G. (1968) A taxonomic study of the family Phytoseiidae (Acari) in South Africa with contributions to the biology of two species. Entomological Memoirs, Department of Agriculture and Water Supply, Republic of South Africa, 18: 1–198.
Vitzthum, H. von (1941) Acarina. In: Bronns, H.G. (Ed.), Klassen und Ordnungen des Tierreichs 5. Akademischer Verlag, Leipzig, Germany, pp. 764–767.
Wainstein, B.A. (1959) New subgenus and species of the genus Phytoseius Ribaga, 1902 (Phytosei-idae: Parasitiformes). Zoologicheskii Zhurnal, 38: 1361–1365.
Wainstein, B.A. (1960) New species and subspecies of the genus Typhlodromus Scheuten (Parasiti-formes, Phytoseiidae) of the USSR fauna. Zoologicheskii Zhurnal, 39: 683–690.
Wainstein, B.A. (1961) New species of mites of the genus Typhlodromus (Parasitiformes: Phytosei-idae) in Georgia. Trudy Instituta Zoologii Akademii Nauk Gruzinskoy SSR, 18: 153–162.
Wainstein, B.A. (1962) Some new predatory mites of the family Phytoseiidae (Parasitiformes) of the USSR fauna. Entomologicheskoe Obozrenie, Russia, 41: 230–240; Entomological Review, 41: 139–146.
Wainstein, B.A. (1972) New species and subgenus of the genus Anthoseius (Parasitiformes, Phytosei-idae). Zoologicheskii Zhurnal, 51: 1477–1482.
Wainstein, B.A. (1975) Predatory mites of the family Phytoseiidae (Parasitiformes) of Yaroslavl Province. Entomologicheskoe Obozrenie, 54(4): 914–922 (In Russian); Entomological Review, 54(4): 138–143.
Wainstein, B.A. (1976) A new tribe of the family Phytoseiidae (Parasitiformes). Zoologicheskii Zhurnal, 55: 696–700.
Wainstein, B.A. & Arutunjan, E.S. (1967) New species of predaceous mites of the genera Typhlodromus Scheuten and Paraseiulus Muma (Parasitiformes, Phytoseiidae). Zoologicheskii Zhurnal, 46: 1764–1770.
Wainstein, B.A. & Arutunjan, E.S. (1968) New species of predaceous mites of the genus Typhlodromus (Parasitiformes: Phytoseiidae). Zoologicheskii Zhurnal, 47: 1240–1244.
Wainstein, B.A. & Vartapetov, S.G. (1973) Predatory mites of the family Phytoseiidae (Parasiti-formes) of Adzharskaya ASSR. Akademiya Nauk Armyanskoy SSR, Biologicheskiy Zhurnal Armenii, 26(2): 102–105.
Walter, D.E. & Krantz, G.W. (2009) Collecting, rearing and preparing specimens. In: Krantz, G.W. & Walter, D.E. (Eds.), A manual of acarology. 3rd ed. Texas Tech University Press, Lubbock, pp. 83–96.
Wu, W.N. & Lan, W.M. (1991) Five new species and one new record of Amblyseius from northwest China (Acari: Mesostigmata: Phytoseiidae). Acta Zootaxonomica Sinica, 16(3): 313–319.
Wu, W.N., Ou, J.F. & Huang, J.L. (2009) Fauna Sinica, Invertebrate Vol. 47, Arachnida, Acari, Phytoseiidae. Beijing: Science Press, 511 pp.
Zaher, M.A. (1986) Predaceous and non‌phytophagous mites (Nile Valley and Delta). Text. Survey and ecological studies on phytophagous, predaceous and soil mites in Egypt. PL 480 Programme USA, Project EG_ARS_30, Grant No. FG_EG_139, Egypt, 567 pp.
Zaher, M.A., Wafa, A.K. & Shehata, K.K. (1969) Life history of the predatory mite Phytoseius plumifer and the effect of nutrition on its biology (Acarina: Phytoseiidae). Entomologia Experimentalis et Applicata, 12: 383–388. DOI: 10.1111/j.1570-7458.1969.tb02534.x
Zannou, I.D. & Hanna, R. (2011) Clarifying the identity of Amblyseius swirskii and Amblyseius rykei (Acari: Phytoseiidae): are they two distinct species or two populations of one species? Experimental and Applied Acarology, 53: 339–347. DOI: 10.1007/s10493-010-9412-6
Zannou, I.D., Moraes, G.J. de, Ueckermann, E.A., Oliveira, A.R., Yaninek, J.S. & Hanna, R. (2006) Phytoseiid mites of the genus Neoseiulus Hughes (Acari: Phytoseiidae) from sub-Saharan Africa. International Journal of Acarology, 32(3): 241–‌276. DOI: 10.1080/01647950608684467
Zhang, Z.-Q. (2003) Mites of Greenhouses: Identification, Biology & Control. CABI Publishing, UK, 244 pp.